Nannaria hardeni

Means, Hennen & Marek, 2021

Curt Harden's Twisted-Claw Millipede

Nannaria hardeni is a of twisted- described in 2021 from the Appalachian Mountains of the eastern United States. It belongs to the Nannaria, a group of forest-dwelling millipedes characterized by males having small, twisted and flattened claws on their legs. The species was named in honor of Curt Harden, a millipede researcher who contributed significantly to the study of Appalachian xystodesmid millipedes. Like other members of its genus, N. hardeni plays a role as a in forest .

Nannaria hardeni (10.3897-zookeys.1030.62544) Figure 29 by Means JC, Hennen DA, Marek PE (2021). Used under a CC BY 4.0 license.Nannaria hardeni (10.3897-zookeys.1030.62544) Figure 28 by Means JC, Hennen DA, Marek PE (2021). Used under a CC BY 4.0 license.

Pronunciation

How to pronounce Nannaria hardeni: /nænˈneɪərɪə ˈhɑːrdɪnaɪ/

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Identification

Males can be identified by their characteristic twisted and flattened on the legs, a of the . -level identification of N. hardeni requires examination of and comparison with closely related species in the Nannaria wilsoni species group. The species was distinguished from through integrative taxonomic analysis including morphological and molecular data.

Images

Appearance

Twisted- in the Nannaria range between 18 and 38 mm in length. They have shiny caramel- to black bodies, often with , red, or orange spots, and white legs. The most distinctive feature is found in males: small, twisted and flattened claws on their legs, which gives the group its . Specific coloration details for N. hardeni have not been separately documented from .

Habitat

Forest floor near streams in the Appalachian Mountains. Like other Nannaria , it is typically found buried under leaf litter, rocks, and logs, often remaining completely beneath the soil surface.

Distribution

Appalachian Mountains of the eastern United States. Specific locality and precise range within the Appalachians have not been separately documented in the provided sources.

Diet

Feeds on decaying leaves and other matter

Life Cycle

Development through typical stages: , with incremental molts adding leg pairs and , . Specific details for N. hardeni not documented separately from .

Behavior

Cryptic : tends to remain buried in soil, sometimes completely beneath the surface. This makes specimens difficult to locate and collect compared to more surface-active relatives.

Ecological Role

: breaks down leaf litter, releasing nutrients into the . Contributes to in forest floor .

Human Relevance

Named in honor of Curt Harden, recognizing his contributions to the study of Appalachian xystodesmid . Part of a 2021 revision that described 17 new Nannaria , advancing understanding of in the Appalachian region.

Similar Taxa

  • Nannaria marianaeAnother described in the same 2021 revision; distinguished by specific and geographic distribution
  • Other Nannaria wilsoni group speciesShare forest floor and twisted- ; require detailed morphological or molecular analysis for separation

More Details

Etymology

Named after Curt Harden in recognition of his significant contributions to the study of xystodesmid in the Appalachian region

Discovery context

Described as part of a comprehensive 2021 revision of the Nannaria wilsoni group based on examination of over 1800 specimens from field collections and museum holdings across 17 US states

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Sources and further reading