Ululodes quadripunctatus
(Burmeister, 1839)
Four-spotted Owlfly
Ululodes quadripunctatus, commonly known as the four-spotted owlfly, is a of owlfly in the and tribe Ululodini. The species is distributed across Central America and North America. Owlflies are neuropteran insects that share characteristics with both and butterflies, possessing large , many-veined wings, and long clubbed .

Pronunciation
How to pronounce Ululodes quadripunctatus: /ˌjuːjəˈloʊdiːz kwɒdrɪˌpʌŋkˈteɪtəs/
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Identification
The Ululodes is distinguished by divided . -level identification of Ululodes quadripunctatus requires examination of specific wing markings; the "four-spotted" refers to characteristic spot patterns on the wings. Detailed identification criteria for this species are not well documented in available sources. Owlflies in general can be distinguished from similar neuropterans by their large bulging eyes, elongated bodies, and clubbed (unlike antlions, which have shorter, less clubbed antennae).
Images
Distribution
Central America and North America. Specific distribution records indicate presence in Middle America and North America according to GBIF data.
Behavior
owlflies in the Ululodes have been observed adopting a cryptic resting posture with the projecting from the perch, resembling a twig. Larvae of Ululodes are , lying on the ground with large trap-jaws held wide open and often camouflaging themselves with sand and debris while waiting for prey. Contact with the jaws triggers them to snap shut, paralyzing prey within minutes.
Ecological Role
As larvae, members of this function as ambush in sandy . The predatory of larvae contributes to soil dynamics.
Similar Taxa
- Ululodes macleayanusAnother in the same with similar and preferences; species-level distinction requires examination of specific wing markings and spot patterns.
- Antlions (Myrmeleontidae)Closely related neuropteran with similar larval predatory habits, but distinguished by shorter, less clubbed in and different larval hunting strategies (pit-building in many ).
More Details
Taxonomic Note
There is taxonomic confusion in available sources regarding the placement of Ululodes. Some sources (GBIF, Catalogue of Life, iNaturalist) place Ululodes in , while others (NCBI, Wikipedia) place it in Myrmeleontidae. The tribe Ululodini is consistently recognized across sources. This reflects ongoing taxonomic revision in the group.