Morphologically-heterogeneous
Guides
Acidocerinae
Acidocerinae is a cosmopolitan subfamily of water scavenger beetles (Hydrophilidae) containing over 500 species in 23-24 genera, making it one of the largest and most morphologically heterogeneous lineages in the family. The subfamily occupies a key phylogenetic position as sister to the largely terrestrial Cylominae+Sphaeridiinae clade. Recent taxonomic revisions (2013, 2021) have substantially restructured generic boundaries, establishing new genera such as Colossochares, Novochares, Aulonochares, Ephydrolithus, and Primocerus while synonymizing former subgenera of Helochares.