Phyllophaga tristis
(Fabricius, 1781)
tristis complex
tristis is a in the , commonly referred to as the "tristis complex." It belongs to a of May/ characterized by soil-dwelling larval development and patterns. The has been studied in relation to allied Phyllophaga forms to understand shared characteristics. Taxonomic placement has been revised, with some sources placing it in the genus Trichesthes.

Pronunciation
How to pronounce Phyllophaga tristis: /ˌfɪl.oʊˈfeɪ.ɡə ˈtrɪs.tɪs/
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Habitat
Soil-dwelling larval stages; terrestrial typical for in the
Distribution
Nearctic region: Canada (British Columbia, Ontario), United States (Colorado, Florida, Georgia, Indiana, Iowa, Louisiana, Maryland, Mississippi, Nebraska, New Mexico, North Dakota, Oregon, South Carolina, Texas, Washington, Wisconsin)
Life Cycle
Involves soil-dwelling larval development typical of ; patterns studied in relation to allied forms
More Details
Taxonomic Uncertainty
GBIF lists tristis as a synonym of Trichesthes tristis, while iNaturalist and NCBI maintain it in Phyllophaga. Catalogue of Life indicates have been synonymized under different (e.g., Phyllophaga tristis amplicornis as synonym of Trichesthes amplicornis), suggesting ongoing taxonomic revision.
Subspecies
Three have been described: tristis amplicornis Reinhard, 1939; Phyllophaga tristis suttonana Reinhard, 1939; and Phyllophaga tristis tristis (Fabricius, 1781). Current taxonomic status of these subspecies is unclear and may require verification against modern revisions.