Cassida piperata
Hope, 1842
Small Tortoise Beetle, tortoise beetle
Cassida piperata is a in the leaf beetle Chrysomelidae. Native to eastern Asia, it has been introduced to North America and Europe, with established in northern Italy since 2021. It serves as a for the Holcotetrastichus rhosaces in Japan.



Pronunciation
How to pronounce Cassida piperata: /kasˈsiː.da piˈpe.ra.ta/
These audio files are automatically generated. While they are not always 100% accurate, they are a good starting point.
Images
Habitat
Anthropized areas including external walls of buildings, log piles, litter, and river banks at elevations of 0-200 m. In its native range, occurs in northeastern Japan.
Distribution
Native to East Palaearctic and Indomalayan Regions; introduced to the United States; established in northern Italy (Veneto: Padova province, Codevigo, Vigonza; Lombardia: Bergamo province, Rogno, Lecco province, Verderio) since 2021.
Seasonality
recorded in May, August, September, October, and November. observed in adults on log piles in November.
Diet
Phytophagous; recorded feeding on Amaranthus retroflexus (redroot pigweed).
Host Associations
- Amaranthus retroflexus - food plantexplicitly recorded as in Italy
Life Cycle
observed in stage. Mating recorded in adults on plants in anthropized areas.
Behavior
Overwinters on log piles. observed mating on plants. Occurs on external walls of buildings and in litter in anthropized .
Ecological Role
for the Holcotetrastichus rhosaces (Walker) (Hymenoptera: Tetrastichinae) in Japan. Potential pest of cultivated plants; non-native with capacity for establishment and spread in Europe based on bioclimatic modelling.
Human Relevance
Non-native with potential to become an agricultural pest. Subject of bioclimatic modelling to assess invasion risk in Europe.
Similar Taxa
- Cassida nebulosaBoth serve as for the same Holcotetrastichus rhosaces in Japan and occur sympatrically in that region.
Sources and further reading
- BugGuide
- Wikipedia
- GBIF taxonomy match
- iNaturalist taxon
- NCBI Taxonomy
- Eaters of thistles: Thistle tortoise beetle, Cassida rubiginosa, and Canada thistle bud weevil, Larinus planus — Bug of the Week
- Northern Ireland, invertebrate finds in review 2023 - Buglife Blog - Buglife
- Little Organisms, Big World: Insect Gut Bacteria Partnerships
- Chrysomelidae | Beetles In The Bush | Page 4
- literature | Beetles In The Bush | Page 5
- Seasonal Variation in a Hymenopterous Parasitoid, Holcotetrastichus rhosaces (Walker) (Hymenoptera: Tetrastichinae), on Its Hosts, Cassida nebulosa L. and C. piperata Hope (Coleoptera: Chrysomelidae), in Sendai, Northeastern Area of Japan
- Cassida piperata Hope, 1843 (Coleoptera, Chrysomelidae), a non-native beetle, new to Italy and Europe.