Helluonini
Hope, 1838
Flat-horned Ground Beetles
Genus Guides
1- Helluomorphoides(Flat-horned Ground Beetles)
Helluonini is a tribe of ground beetles within the Anthiinae, containing more than 20 and approximately 190 described . Members are distributed across tropical and subtropical regions including the Oriental, Ethiopian, and Palaearctic realms. The tribe includes medium-sized beetles (8–17 mm) with characteristic elongate bodies, coarse punctation, and . The genus Macrocheilus, the most speciose in the tribe with 56 species, exemplifies the group's morphological diversity across Africa, Asia, and Madagascar.



Pronunciation
How to pronounce Helluonini: /hɛˌljuːoʊˈnaɪniː/
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Identification
Helluonini can be distinguished from other ground beetle tribes by the combination of: two supraorbital setae on the ; deeply emarginated with a long, slender tooth; and fully functional wings. Members of the Macrocheilus show no external . Body form is elongate with coarse surface punctation and . Size ranges from 8.0 to 17.0 mm in the best-studied genus.
Images
Distribution
Tropical and subtropical regions across three biogeographic realms: Oriental Region (China, Vietnam, Laos, Cambodia, Myanmar, India, Sri Lanka, Philippines, Malaysia, Indonesia); Ethiopian Region (Ethiopia, Tanzania, Uganda, Senegal, Guinea, Guinea-Bissau, Sierra Leone, Togo, Nigeria, Central Africa, Cameroon, Gabon, Congo, Zimbabwe, Angola, South Africa, Madagascar); and Palaearctic Region (Pakistan, Syria, Asia Minor).
Similar Taxa
- Other Anthiinae tribesHelluonini shares the Anthiinae with other tribes; distinguished by the combination of two supraorbital setae, emarginated with long slender tooth, and functional wings.
More Details
Taxonomic history
Authored by Hope in 1838. The tribe has undergone significant taxonomic revision, particularly for Oriental of Macrocheilus.
Species diversity
The Macrocheilus comprises 56 total: 35 Ethiopian, 19 Oriental, and 2 Palaearctic. The tribe as a whole exceeds 190 described species across more than 20 genera.