Trechitae
Guides
Horologion hubbardi
Horologion hubbardi is a newly described species of carabid beetle from Bath County, Virginia. It represents only the second known species in the genus Horologion, which was previously known solely from a single holotype collected in West Virginia in 1931. DNA sequence data and morphological analysis place the genus in the supertribe Trechitae, as sister to the Gondwanan tribe Bembidarenini. The species is considered a relict lineage with no close relatives known in the Northern Hemisphere.
Horologionini
Horologionini is a relict tribe of carabid beetles (Coleoptera: Carabidae: Trechinae) containing the enigmatic genus Horologion. The tribe was newly recognized in 2024 based on DNA sequence data and morphological analysis. Phylogenetic analysis places Horologionini in the supertribe Trechitae as sister to the Gondwanan tribe Bembidarenini, rejecting previous hypotheses that placed the genus near Anillini, Tachyini, Trechini, Patrobini, or Psydrini. The tribe has no close relatives in the Northern Hemisphere and represents an important component of Appalachian biodiversity.
Phrypeus
Phrypeus is a monotypic genus of ground beetles (Carabidae: Trechinae: Sinozolini) containing the single species Phrypeus rickseckeri. The genus was described by Casey in 1924 and is considered taxonomically isolated within the supertribe Trechitae. Its phylogenetic relationships remain obscure; larval morphology studies have failed to link it definitively with any other tribe. The first-instar larva has been described and illustrated.