Extreme-rarity
Guides
Horologionini
Horologionini is a relict tribe of carabid beetles (Coleoptera: Carabidae: Trechinae) containing the enigmatic genus Horologion. The tribe was newly recognized in 2024 based on DNA sequence data and morphological analysis. Phylogenetic analysis places Horologionini in the supertribe Trechitae as sister to the Gondwanan tribe Bembidarenini, rejecting previous hypotheses that placed the genus near Anillini, Tachyini, Trechini, Patrobini, or Psydrini. The tribe has no close relatives in the Northern Hemisphere and represents an important component of Appalachian biodiversity.