Lithobiomorpha
Stone Centipedes
Family Guides
2- Henicopidae
- Lithobiidae(stone centipedes)
Lithobiomorpha, commonly known as stone centipedes, are an order of anamorphic characterized by a mature segment count of 15 trunk segments. They lack , possessing instead simple ocelli or no in some subterranean . The order includes two : Henicopidae and Lithobiidae. Members are primarily found in forest litter and soil across temperate regions worldwide, with significant diversity in China and Europe.



Pronunciation
How to pronounce Lithobiomorpha: //ˌlaɪθoʊˌbaɪoʊˈmɔːrfə//
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Identification
Distinguished from Scutigeromorpha by shorter and legs. Differentiated from other orders by the combination of 15 mature trunk segments, simple ocelli (not ), and paired lateral . Antennal segmentation (15 or 17 segments) serves as a reliable character within the Anopsobiinae . Cave-dwelling may be eyeless, requiring careful examination of other morphological traits.
Images
Appearance
possess 15 trunk segments with a single pair of legs per segment. Each leg-bearing segment has a separate tergite, alternating in length except for paired long tergites on segments 7 and 8. and legs are relatively short compared to Scutigeromorpha. are simple ocelli or groups of ocelli; some cave-dwelling lack eyes entirely. are paired and positioned laterally.
Habitat
Primarily inhabits forest litter and soil. Frequently found under stones and in leaf litter in conifer and larch forests. Some occur in caves, though most cave records represent surface-dwelling species rather than obligate troglobites. Moist microhabitats preferred; recorded from elevations ranging from 950 m to 1400 m in mountainous regions of China.
Distribution
Widespread across temperate regions of the Northern Hemisphere. Documented from China (68 , 44 ), Europe including Slovenia (21 species in caves), former USSR territories, Australasia (Australia, New Zealand, New Caledonia), South Africa, and southern South America. Records from Kazakhstan, northwestern China (Xinjiang), and the Qinghai-Tibet Plateau.
Diet
Predominantly , feeding on insects and other small . Has been observed to feed on leaf litter and associated small animals, indicating broader dietary habits than previously assumed. Some occasionally consume slugs and worms.
Life Cycle
Anamorphic development: juveniles hatch with fewer segments and leg pairs, adding segments and legs through successive until reaching the mature count of 15 trunk segments. Specific details on number, developmental duration, and longevity vary by and remain incompletely documented.
Behavior
that actively hunt in soil and leaf litter. Some secrete sticky fluids to immobilize prey. Fast-moving compared to millipedes. Bite is painful and venomous, comparable in intensity to a sting.
Ecological Role
of insects and other soil ; play an essential role in regulating densities of soil invertebrate . Contribute to nutrient cycling through and, in some , consumption of leaf litter.
Human Relevance
Generally beneficial in gardens and landscapes as of pest insects. Bites can cause pain comparable to stings; handling with bare hands is not recommended. Occasionally enter homes; can be deterred by maintaining bare ground barriers between mulch and foundations and ensuring proper door seals.
Similar Taxa
- ScutigeromorphaHouse centipedes and relatives possess and extremely long legs and , contrasting with Lithobiomorpha's simple ocelli and shorter appendages.
- Julida (millipedes)Millipedes have two pairs of legs per body segment, are generally slower, and are rather than ; Lithobiomorpha have one pair of legs per segment and are active predators.
More Details
Taxonomic Families
The order comprises two : Henicopidae and Lithobiidae. Chinese fauna includes 15 /subgenera, with three genera (Pterygotergum, Hedinobius, and one additional) to China.
Cave Occurrence
In Slovenian caves, 21 lithobiomorph have been recorded, but only three (Lithobius stygius, L. zveri, and Eupolybothrus obrovensis) are considered exclusively subterranean; most records represent surface species.
Research Gaps
The myriapod fauna of China remains poorly known, with stone centipedes receiving particularly limited attention. The role and importance of lithobiomorphs in subterranean remain to be studied.
Sources and further reading
- BugGuide
- Wikipedia
- GBIF taxonomy match
- iNaturalist taxon
- NCBI Taxonomy
- Catalogue of Life
- Pensoft Editorial Team | Blog - Part 84
- Life and death down under - Garden millipedes (Order Julida) and stone centipedes (Genus Lithobius — Bug of the Week
- An annotated checklist of Lithobiomorpha of China
- Molecular phylogeny of Australasian anopsobiine centipedes (Chilopoda : Lithobiomorpha)
- Distribution patterns of lithobiomorpha on the territory of the USSR
- The overview of lithobiomorph centipedes (Chilopoda, Lithobiomorpha) from caves of Slovenia
- Bacterial diversity associated with the brown stone centipede, Lithobius forficatus (Chilopoda, Lithobiomorpha)
- Новые данные о Australobius magnus (Trotzina, 1894) (Chilopoda: Lithobiomorpha: Lithobiidae из Южного Казахстана
- A review of the Chinese species of Bothropolys Wood, 1862 (Chilopoda: Lithobiomorpha: Lithobiidae)
- A new species of the genus Australobius Chamberlin, 1920 (Lithobiomorpha, Lithobiidae) from southwestern China.