Anemadini
Hatch, 1927
Anemadini is a tribe of small within the Cholevinae ( ). The tribe comprises at least 30 and more than 70 described . Members exhibit diverse ecological , including forest-dwelling carrion feeders, cave inhabitants, and myrmecophiles living in nests. Some lineages show morphological modifications associated with subterranean life, such as reduced or absent .
Pronunciation
How to pronounce Anemadini: //əˌnɛməˈdaɪnaɪ//
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Identification
Anemadini can be distinguished from other tribes in Cholevinae by a involving the female genital annulus. Within the tribe, subtribes Eocatopina and Nemadina can be differentiated; Eocatopina includes the Neoeocatops in North America, while Nemadina includes Nemadus and related genera. in the Anemadus smetanai group show gradual morphological modifications linked to endogean life, including microphthalmy and anophthalmy.
Habitat
vary by lineage. Forest-dwelling occur in woodland environments. Some species inhabit caves. The Anemadus smetanai species group occupies subterranean, high-altitude in China. Myrmecophilous species occupy nests of formicine .
Distribution
The tribe has a broad distribution including North America, the Western Mediterranean (Iberian Peninsula, North Africa, Sicily, continental Italy), and the Palaearctic region (China: Sichuan, Yunnan, Xizang). The Speonemadus is restricted to the Western Mediterranean. The genus Anemadus occurs in the Palaearctic region.
Diet
Carrion and other decomposing materials. This feeding habit has been documented specifically for Nemadus in North America.
Host Associations
- Formica - myrmecophileformicine nests
- Camponotus - myrmecophileformicine nests
Behavior
: some occupy nests of formicine , primarily Formica and . Subterranean species show morphological to endogean life.
Ecological Role
. feed on carrion and decaying materials, contributing to in forest and subterranean .
Similar Taxa
- other Cholevinae tribesAnemadini is distinguished from other tribes in the by the female genital annulus ; other tribes lack this character and typically show different ecological associations or geographic distributions
More Details
Systematic subdivisions
The tribe contains at least two subtribes: Eocatopina (containing Neoeocatops in North America) and Nemadina (containing Nemadus and related ). The subtribe Eocatopina was newly recorded from North America in 2005.
Evolutionary history
The Speonemadus diversified from the late Miocene (Tortonian), with cladogenetic events linked to the isolation of landmasses between Iberia and North Africa, including the opening of the Strait of Gibraltar at the end of the Messinian.
Sources and further reading
- BugGuide
- Wikipedia
- iNaturalist taxon
- Catalogue of Life
- Subterranean species of Anemadus Reitter: systematics, phylogeny and evolution of the Chinese "Anemadus smetanai" species group (Coleoptera: Leiodidae: Cholevinae: Anemadini)
- A molecular phylogeny of Speonemadus Jeannel, 1922 with description of two new species from Morocco (Coleoptera: Leiodidae: Cholevinae: Anemadini)
- Systematics, distributions, and bionomics of the Neoeocatops gen. nov. and Nemadus of North America (Coleoptera: Leiodidae: Cholevinae: Anemadini)