Philodromidae
Thorell, 1870
Running Crab Spiders, Philodromid Crab Spiders
Genus Guides
7- Apollophanes(running crab spiders)
- Ebo(running crab spider)
- Philodromus(Running Crab Spiders)
- Rhysodromus
- Thanatus(false crab spiders)
- Tibellus(Slender Crab Spiders)
- Titanebo
Philodromidae is a of araneomorph spiders comprising over 530 across 31 . Members are commonly called running crab spiders or philodromid crab spiders due to their crab-like appearance and sideways-oriented legs. Unlike true crab spiders (Thomisidae), philodromids do not construct webs for prey capture but instead hunt by ambush. They are primarily associated with vegetation, where they occur on stems, leaves, and tree trunks.



Pronunciation
How to pronounce Philodromidae: /fɪloʊˈdroʊmɪˌdeɪ/
These audio files are automatically generated. While they are not always 100% accurate, they are a good starting point.
Identification
Philodromids possess laterigrade legs—oriented sideways like those of crabs—with the first two pairs projecting forward. The second leg pair is often longer than the first, most dramatically in the Ebo where it may be twice as long. are arranged in two curved rows with the row wider than the . Scopulae (adhesive hairs) are present only at the tarsal tips, unlike in Sparassidae. Body coloration is typically cream to light brown with faint longitudinal stripes. The overall body form is flattened.
Images
Habitat
Primarily associated with vegetation including grasses, shrubs, tree trunks, low herbage, and undergrowth. Some occur specifically on deciduous trees, others on conifers. A small number inhabit desert environments.
Distribution
Worldwide distribution. Documented from North America (including southern California to western Texas, Florida, and the Caribbean), Europe, Asia (including South Korea, China, Japan), Africa (Afrotropical region), and the Neotropics.
Seasonality
and immatures overwinter in temperate regions, becoming active in early spring when temperatures rise. Peak activity observed from March through May in North American temperate zones.
Diet
Active that hunt by ambush. Prey consists of insects and other small arthropods captured directly without web use.
Life Cycle
Females construct flattened sacs which they guard until spiderlings emerge. occurs as or immatures in leaf litter, grass tussocks, or other sheltered microhabitats. Spiders contain glycerol and specialized proteins in that reduce freezing point, enabling cold .
Behavior
Non-web-building hunters that rely on speed and ambush rather than silk traps. When disturbed, they can sprint with surprising speed. Some are known to enter buildings; the house crab spider (Thanatus) commonly captures flies on and in structures.
Ecological Role
of insects and small arthropods in vegetation . Serve as prey for spider wasps (Pompilidae), including Priocnemis minorata which has been documented parasitizing Thanatus.
Human Relevance
Not dangerously venomous to humans. Occasionally encountered indoors where they may capture pest flies. Frequently mistaken for true crab spiders (Thomisidae) or, in the southwestern U.S. and Florida, for the unrelated 'flatties' (Selenopidae).
Similar Taxa
- ThomisidaeTrue crab spiders share laterigrade legs and crab-like appearance, but differ in having markedly unequal leg sizes with first two pairs much longer and stouter, and in building no sacs (guarding eggs under silk instead).
- Selenopidae'Flatties' share extreme flattening and laterigrade legs, but have six in one row (vs. two curved rows in Philodromidae), legs increasingly longer from front to back (vs. second pair longest in Philodromidae), and a notched rear sternum edge.
- SparassidaeGiant crab spiders may appear similar but have scopulae covering entire rather than just tips, and different arrangement.
Sources and further reading
- BugGuide
- Wikipedia
- GBIF taxonomy match
- iNaturalist taxon
- NCBI Taxonomy
- Catalogue of Life
- Bug Eric: Spider Sunday: Spring Spiders
- Bug Eric: Spider Sunday: Flatties
- Bug Eric: Spider Sunday: Slender Crab Spiders (and their look-a-likes)
- Bug Eric: Priocnemis minorata
- Bug Eric: A Nursery Web Spider, Pisaurina dubia
- Bug Eric: Shower Spiders
- Taxonomic revision of the genus Gephyrellula Strand, 1932 (Arachnida, Araneae, Philodromidae)
- A revision of the Afrotropical species of the genus<i> Tibellus </i>Simon (Araneae: Philodromidae)
- Philodromus uljin sp. nov., a new running crab spider (Araneae, Philodromidae) from South Korea.
- On small huntsman spiders (Araneae, Philodromidae) occurring in Guizhou and Hubei provinces, China.