Condition-dependent-trait
Guides
Gnatocerus cornutus
Broadhorned Flour Beetle, Broad-horned Flour Beetle
Gnatocerus cornutus is a globally distributed stored-product pest in the darkling beetle family Tenebrionidae. Males possess enlarged mandibles used as weapons in intrasexual combat, representing a well-studied model for sexual selection and condition-dependent trait expression. The species exhibits striking sexual dimorphism and intra-sexual dimorphism in males, with weapon size strongly influenced by larval nutrition.