Tortricoidea
Latreille, 1803
Tortricid Leafroller Moths and Allies
Family Guides
1- Tortricidae(tortrix moths)
is a superfamily of Lepidoptera containing the single Tortricidae, commonly known as tortrix moths or leafroller moths. This is one of the largest superfamilies of , with over 10,350 described . Members are characterized by their typical resting posture with wings folded back, producing a rounded profile. Many species are economically important agricultural and forestry pests.



Pronunciation
How to pronounce Tortricoidea: //tɔrˌtrɪkoʊˈɪdiə//
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Identification
rest with wings folded tightly over the , creating a distinctive bell-shaped or rounded silhouette. The forewings are typically broad and somewhat rectangular, often with banded or mottled patterns. Hindwings are membranous and narrower, folded beneath the forewings at rest. are . Larvae are often concealed within rolled or tied leaves, or bore into fruits, seeds, or buds.
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Habitat
Found in diverse terrestrial including forests, orchards, agricultural fields, and gardens. Many are associated with woody plants, particularly conifers and deciduous trees.
Distribution
distribution with occurring worldwide. The Tortricidae alone has been documented from India (487 species), Europe, North America, and the Murcia Region of Spain (107 species), among other regions.
Diet
Larvae feed on a wide variety of plant materials including leaves, fruits, seeds, buds, and shoots of numerous plant . Many are on particular plants. Specific dietary preferences vary extensively among species.
Life Cycle
Complete with , larval, pupal, and stages. Larval development typically occurs within protected microhabitats such as rolled leaves, tied foliage, or bored plant tissues. Some overwinter as larvae, others as eggs. Voltinism (number of per year) varies by species and climate, with , , and multivoltine species documented.
Behavior
Larvae are the primary feeding stage and often construct shelters by rolling or tying leaves with silk. Many are known as "leafrollers" due to this . are typically and attracted to light.
Ecological Role
Larvae function as herbivores, with many acting as significant defoliators of trees and shrubs. They serve as for diverse complexes including ichneumonid wasps, braconid wasps, and other parasitic insects. Some species are considered keystone pests in forest and agricultural .
Human Relevance
Many are major agricultural and forestry pests causing substantial economic damage to crops including apples, pears, stone fruits, berries, and conifer forests. The superfamily includes notable pests such as codling moth and various budmoths. Some species have been targets for programs.
Similar Taxa
- PyraloideaBoth are large superfamilies of with diverse larval feeding habits; can be distinguished by the characteristic resting posture with wings folded tightly back creating a rounded profile, versus the more triangular wing posture typical of many pyraloids.
- CossoideaSome wood-boring larvae may appear similar; of are generally smaller with more rounded wing profiles and lack the elongated bodies of many cossoid .
Sources and further reading
- BugGuide
- iNaturalist taxon
- NCBI Taxonomy
- Catalogue of Life
- A Gathering of Lepidopterists: Who's Who of the Butterfly, Moth Experts | Bug Squad
- Bug Eric: October 2010
- Tortricoidea: Tortricidae: Olethreutinae, Tortricinae, undetermined subfamily
- Catalogue of Tortricidae Latreille, 1802 (Lepidoptera: Tortricoidea) of India
- Observations on the parasitoid complexes of budmoths (Lepidoptera: Tortricoidea) on larch in Europe
- Parasitoid Guilds, Life-Styles, and Host Ranges in the Parasitoid Complexes of Tortricoid Hosts (Lepidoptera: Tortricoidea)
- An annotated checklist of the Tortricidae of the region of Murcia (Spain) with new records, distribution and biological data (Lepidoptera, Tortricoidea)
- Figure 2 from: Garre MJ, Girdley J, Guerrero J, Rubio RM, Ortiz AS (2025) An annotated checklist of the Tortricidae of the region of Murcia (Spain) with new records, distribution and biological data (Lepidoptera, Tortricoidea). Biodiversity Data Journal 13: e150786. https://doi.org/10.3897/BDJ.13.e150786
- Figure 1 from: Garre MJ, Girdley J, Guerrero J, Rubio RM, Ortiz AS (2025) An annotated checklist of the Tortricidae of the region of Murcia (Spain) with new records, distribution and biological data (Lepidoptera, Tortricoidea). Biodiversity Data Journal 13: e150786. https://doi.org/10.3897/BDJ.13.e150786