Tomoderinae
Bonadona, 1961
Genus Guides
1Tomoderinae is a of beetles within the Anthicidae (Coleoptera: Tenebrionoidea), established by Bonadona in 1961. The subfamily includes the Macrotomoderus Pic, 1901, which is the most -rich and well-documented genus, with numerous species described from the Palaearctic region, particularly China, Taiwan, and the Japanese Archipelago. Species of Macrotomoderus are predominantly flightless () and exhibit high-altitude distribution patterns in montane regions of continental China.

Pronunciation
How to pronounce Tomoderinae: //ˌtoʊmoʊdəˈraɪniː//
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Identification
Members of Tomoderinae can be distinguished from other Anthicidae by morphological characters detailed in taxonomic revisions, including features of the male genitalia and body proportions. Within the subfamily, Macrotomoderus are identified using keys based on habitus and male genitalia structure. Some species exhibit distinctive traits, such as the dense clump of curled setae at the base in male M. hirsutus, which forms a subconical bristle concealing the pronotal margin.
Images
Habitat
Montane regions at moderate to high elevations. Specific elevation records for Macrotomoderus include 1700–1750 m a.s.l., 1900 m a.s.l., and 2800–3200 m a.s.l. in the Daba Mountains and adjacent ranges of central China.
Distribution
Palaearctic region, with documented occurrence in continental China, Taiwan, and the Japanese Archipelago. Within China, have been recorded from montane areas in Sichuan, Hubei, Shaanxi, and adjacent provinces. Distribution follows altitudinal gradients, with species occupying distinct elevation bands.
Behavior
(flightless) condition has been documented in multiple Macrotomoderus . No other behavioral data are available.
More Details
Taxonomic focus
Available literature on Tomoderinae consists primarily of taxonomic revisions describing new of Macrotomoderus and providing identification keys. Ecological, biological, and behavioral data remain largely unstudied for this .
Biogeography
The altitudinal gradient of Macrotomoderus distribution in continental China has been discussed in taxonomic literature, suggesting elevation-based segregation in montane .
Sources and further reading
- BugGuide
- iNaturalist taxon
- Catalogue of Life
- Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species
- Revision of the Tomoderinae (Coleoptera, Anthicidae). Part V. Three new Macrotomoderus Pic, 1901 from continental China and an updated key to the Palaearctic species.