Empidoidea
Dance Flies, Long-legged Flies, and Allies
is a large superfamily of () comprising approximately 10,000 described across all continents except Antarctica. As the sister to Muscomorpha (), these two groups are sometimes united in the unranked taxon . The superfamily includes several well-known : (dagger and ), (), and Hybotidae (). The group has existed since the period and has been subject to extensive phylogenetic revision, particularly following molecular studies in 2018 that reshaped relationships among constituent families.



Pronunciation
How to pronounce Empidoidea: /ˌɛmpɪˈdɔɪdiːə/
These audio files are automatically generated. While they are not always 100% accurate, they are a good starting point.
Identification
can be distinguished from other by the combination of large that often cover most of the surface, and that frequently includes a well-developed and/or a small but distinct . Members lack the ptilinal fissure characteristic of . are typically aristate with three . The superfamily is morphologically diverse, with ranging from the elongate-legged to the more compact Hybotidae.
Images
Habitat
The majority of are associated with moist, temperate . Many have involving freshwater environments. Specific habitat preferences vary considerably among , with some groups found in agricultural systems such as winter wheat fields.
Distribution
occurs on all continents except Antarctica, with representation in both Northern and Southern Hemispheres including Chile, Tibet, and Nearctic regions. The superfamily has a global distribution spanning diverse biogeographic regions.
Diet
The majority of are predatory, feeding on other small . Hybotidae in particular have been documented as predatory in agricultural systems.
Behavior
Several exhibit elaborate mating rituals that have attracted interest from ethologists. Many species have involving freshwater . Specific behavioral details vary widely among and remain poorly documented for many groups.
Ecological Role
As predominantly predatory , function as of other small in their . Hybotidae have been studied as components of predator in agricultural .
Human Relevance
has significant scientific importance for studies of dipteran and evolution, with the group serving as a lineage for understanding relationships among higher . Some are subjects of ethological research due to their complex mating behaviors. The group is taxonomically active with ongoing species description and revision.
Similar Taxa
- Cyclorrhapha (Muscomorpha)Sister to ; distinguished by presence of ptilinal fissure used for from , and generally different patterns. Empidoidea lacks the and associated .
- AsilomorphaAnother major clade of predatory ; Asilomorpha includes () and related , which differ in body plan, , and typically have a more , stout-bodied appearance compared to the generally more slender .
Sources and further reading
- BugGuide
- Wikipedia
- iNaturalist taxon
- NCBI Taxonomy
- Super Crop Challenge #4 | Beetles In The Bush
- How to Describe a New Species: Getting into the “Spuh Nov” Club
- A catalog of the Brachystomatidae of Chile (Diptera: Empidoidea)
- Heleodromia Haliday newly found in Tibet with description of one new species (Diptera: Empidoidea: Trichopezinae)
- World revision of Iteaphila with unbranched radial vein (Diptera: Empidoidea: Iteaphilidae)
- Review of three little-known monotypic empidoid genera (Diptera: Empidoidea: Brachystomatidae), assigned to Trichopezinae
- Community Composition of Predatory Hybotidae (Diptera: Empidoidea) in Winter Wheat Management Systems.